During the Devonian the sedimentation on the continental shelves of Ardenne Massif and Boulonnais has changed from a mixed siliciclastic-carbonate ramp (Eifelian), through a carbonate barrier reef (Givetian) and then to a detritic influx with local mud-mounds (Frasnian). Here we analysed the faunistic dynamics of the trilobite associations through the changing environment. We used multivariate analyses (clustering and ordering) to discriminate the trilobite associations within 67 different samples. Three previously known communities and one new were recognised: the Eifelian Mixed association, the Givetian Dechenella association and the two Frasnian Bradocryphaeus and Scutellum-Goldius associations. These trilobite faunas present a progressive ecological specialisation. The Mixed association occurs both in the ramp or carbonated (local reef developed on the ramp) facies without any significant difference in its composition. The Dechenella fauna occurs preferentially close to barrier reefs, but can also survive during short periods of detrital input. The two Frasnian communities show a strong relationship with their environment. The Scutellum-Goldius association is only found in reef systems, whereas the Bradocryphaeus flourishes exclusively in lateral facies. ; Fil: Bignon, Arnaud Marcel Jacques. Consejo Nacional de Investigaciones Científicas y Técnicas. Centro Científico Tecnológico Conicet - Córdoba. Centro de Investigaciones en Ciencias de la Tierra. Universidad Nacional de Córdoba. Facultad de Ciencias Exactas Físicas y Naturales. Centro de Investigaciones en Ciencias de la Tierra; Argentina ; Fil: Crônier, Catherine. Universite Lille 3; Francia